Ronald Joseph Livingston was born on June 5, 1967 in Cedar Rapids, Iowa, to Linda (Rinas), a Lutheran pastor, and Kurt Livingston, an aerospace engineer. He has three siblings, Nick, John Livingston, also an actor, and Jennifer Livingston, a TV news personality at CBS/WKBT in La Crosse, Wisconsin. He is of German, and smaller amounts of Welsh, Scottish, and English, ancestry.